Use scientific expression translate the sentences随后他们还发现17LamD内脂环上的羰基、C-8位和C-20位的羟基分别能能与Topo I 结合口袋中的3个重要结合位点Arg 364 Asn 722 Glu 356 产生相互作用形成三元复合物。

Subsequently, they also discovered that the carbonyl group on the LamD inner lipid ring and the hydroxyl groups on C-8 and C-20 can interact with the three important binding sites (Arg 364, Asn 722, Glu 356) in the pocket of Topo I, forming a ternary complex.