Use a more scientific expression translate these sentences：随后他们还发现17LamD内脂环上的羰基、C-8位和C-20位的羟基分别能能与Topo I 结合口袋中的3个重要结合位点Arg 364 Asn 722 Glu 356 产生相互作用形成三元复合物。

Subsequently, they also observed that the carbonyl group on the LamD inner lipid ring, as well as the hydroxyl groups on C-8 and C-20, could interact with three critical binding sites (Arg 364, Asn 722, Glu 356) in the Topo I binding pocket, forming a ternary complex.